The game features nine dungeons—large, contained areas where Link battles enemies, collects items, and solves puzzles. Link navigates these dungeons and fights a boss at the end in order to obtain an item or otherwise advance the plot. The dungeons are connected by a large overworld, across which Link can travel on foot; on his horse, Epona; or by teleporting.
When Link enters the Twilight Realm, the void that corrupts parts of Hyrule, he transforms into a wolf. He is eventually able to transform between his Hylian and wolf forms at will. As a wolf, Link loses the ability to use his sword, shield, or any secondary items; he instead attacks by biting, and defends primarily by dodging attacks. However, “Wolf Link” gains several key advantages in return—he moves faster than he does as a human (though riding Epona is still faster) and digs holes to create new passages and uncover buried items, and has improved senses, including the ability to follow scent trails. He also carries Midna, a small imp-like creature who gives him hints, uses an energy field to attack enemies, helps him jump long distances, and eventually allows Link to “warp” to any of several preset locations throughout the overworld. Using Link’s wolf senses, the player can see and listen to the wandering spirits of those affected by the Twilight, as well as hunt for enemy ghosts named Poes.
The artificial intelligence (AI) of enemies in Twilight Princess is more advanced than that of enemies in The Wind Waker. Enemies react to defeated companions and to arrows or slingshot pellets that pass by, and can detect Link from a greater distance than was possible in previous games.